Editorial Board

EDITOR IN CHIEF (2017)

Rozanne M. Sandri-Goldin

University of California, Irvine

EDITORS

Lawrence Banks, Editor (2020) International Centre for Genetic Engineering and Biotechnology

Karen L. Beemon, Editor (2017) Johns Hopkins University

Byron W. Caughey, Editor (2018) Rocky Mountain Laboratories

Carolyn Coyne, Minireview Editor (2020) University of Pittsburgh School of Medicine

Terence S. Dermody, Editor (2019) University of Pittsburgh School of Medicine

Michael S. Diamond, Editor (2018) Washington University School of Medicine

Klaus Frueh, Editor (2017) Oregon Health & Science University

Adolfo García-Sastre, Editor (2020) Icahn School of Medicine at Mount Sinai

Britt Glaunsinger, Minireview Editor (2020) University of California, Berkeley

Jae U. Jung, Editor (2020) University of Southern California

Frank Kirchhoff, Editor (2019) Ulm University Medical Center

Richard M. Longnecker, Editor (2021) Northwestern University

Susana López, Editor (2018) Instituto de Biotecnologia/UNAM

Douglas S. Lyles, Editor (2017) Wake Forest University

Grant McFadden, Editor (2017) Arizona State University

J.-H. James Ou, Editor (2019) University of Southern California

Stanley Perlman, Editor (2017) University of Iowa

Julie Pfeiffer, Editor (2021) University of Texas Southwestern Medical Center

Theodore C. Pierson, Minireview Editor (2020) National Institute of Allergy and Infectious Diseases

Susan R. Ross, Editor (2017) University of Illinois at Chicago

Stacey Schultz-Cherry, Cover Editor (2020) St. Jude Children's Research Hospital

Guido Silvestri, Editor (2018) Emory University

Anne Simon, Editor (2017) University of Maryland

Wesley I. Sundquist, Editor (2019) University of Utah

Bryan Williams, Editor (2018) Hudson Institute of Medical Research

ASSOCIATE EDITORS

Michaela Gack, Associate Editor (2018) University of Chicago

Felicia Goodrum, Associate Editor (2019) University of Arizona

Joanna L. Shisler, Associate Editor (2019) University of Illinois

Christopher S. Sullivan, Associate Editor (2019) University of Texas at Austin

Billy Tsai, Associate Editor (2019) University of Michigan Medical School

Thomas E. Shenk, Chairman, Journals Board

Stefano Bertuzzi, Chief Executive Officer

Barbara M. Goldman, Director, Journals

Julie Nachman, Production Editor

Karen Esplin, Assistant Production Editor

Diane Smith, Editorial Coordinator

EDITORIAL BOARD

Allison Abendroth ('19)

Jin-Hyun Ahn ('19)

Tero Ahola ('18)

Chris Aiken ('18)

Randy Albrecht ('18)

Lorraine Albritton ('19)

José Alcami ('18)

Elliot J. Androphy ('17)

Cristian Apetrei ('19)

Carlos F. Arias ('18)

Michelle M. Arnold ('19)

James Arthos ('18)

Ann Arvin ('18)

Aravind Asokan ('17)

Walter J. Atwood ('19)

Steve Bachenheimer ('18)

Joel Baines ('18)

Susan C. Baker ('17)

Bruce W. Banfield ('19)

Eric Barklis ('17)

Dan Barouch ('18)

Alan Barrett ('18)

Peter A. Barry ('19)

Ralf Bartenschlager ('17)

David J. Barton ('17)

Jason C. Bartz ('17)

Chris Basler ('18)

George Belov ('18)

Jeffrey M. Bergelson ('17)

Cornelia Bergmann ('19)

Ben Berkhout ('18)

Sonja Best ('18)

Joel Blankson ('18)

Gary W. Blissard ('19)

David C. Bloom ('19)

Karl W. Boehme ('19)

Kathleen Boris-Lawrie ('18)

Michel Brahic ('18)

Jason M. Brenchley ('17)

Wade Bresnahan ('17)

Margo A. Brinton ('17)

William Britt ('18)

Chris Broder ('18)

Jay Brown ('17)

Wolfram Brune ('19)

Michael Buchmeier ('18)

Christopher B. Buck ('17)

Alexander Bukreyev ('18)

Andrew Byrnes ('19)

Craig Cameron ('18)

Gabriella Campadelli-Fiume ('19)

Massimo Caputi ('19)

Roberto Cattaneo ('18)

Ann Chahroudi ('19)

Bala Chandran ('17)

Wen Chang ('17)

Yu-Chan Chao ('17)

Bruce W. Chesebro ('18)

Gregory Chinchar ('18)

Nicolas Chomont ('19)

François Clavel ('18)

Rollie Clem ('18)

Jeffrey I. Cohen ('19)

Anamaris Colberg-Poley ('18)

Ronald Collman ('18)

Richard C. Condit ('19)

John H. Connor ('19)

Karl-Klaus Conzelmann ('19)

Barbara S. Coulson ('19)

Bryan Cullen ('19)

Blossom Damania ('19)

Pranav Danthi ('19)

Miles Davenport ('17)

David J. Davido ('17)

Andrew J. Davison ('19)

Victor DeFilippis ('18)

Juan Carlos de la Torre ('17)

Neal A. DeLuca ('18)

Mark R. Denison ('17)

Cynthia A. Derdeyn ('19)

Prashant Desai ('18)

Aravinda M. de Silva ('19)

Ronald Desrosiers ('18)

Ulrich Desselberger ('18)

Paul Digard ('18)

Daniel DiMaio ('19)

Dimiter Dimitrov ('18)

Shou-Wei Ding ('17)

Dirk Dittmer ('19)

Valerian Dolja ('17)

Esteban Domingo ('18)

Margherita Doria ('18)

Philip R. Dormitzer ('19)

Christian Drosten ('18)

Jean D. Dubuisson ('17)

Jaquelin Dudley ('18)

Sioban Duffy ('17)

Lynn B. Dustin ('19)

Rebecca E. Dutch ('18)

Gregory Ebel ('19)

John H. Elder ('19)

Alan Engelman ('18)

Lynn W. Enquist ('19)

David T. Evans ('19)

Matthew J. Evans ('19)

Oliver T. Fackler ('19)

Erik Falck-Pedersen ('19)

Bentley Fane ('18)

Paul J. Farrell ('19)

Michael Farzan ('18)

Herman W. Favoreel ('19)

Heinrich U. Feldmann ('17)

Pinghui Feng ('18)

Zongdi Feng ('19)

Ana Fernandez-Sesma ('18)

Jane Flint ('17)

Steven Foung ('19)

Manuel Franco ('18)

Lori Frappier ('19)

Eric O. Freed ('17)

Matthew B. Frieman ('17)

Robert S. Fujinami ('17)

Thomas Gallagher ('17)

Shou-Jiang Gao ('17)

Robert Garcea ('18)

Fernando Garcia-Arenal ('18)

Adam P. Geballe ('19)

Chou-Zen Giam ('18)

Paul Goepfert ('18)

Tatyana V. Golovkina ('17)

Felicia Goodrum ('19)

Alexander Gorbalenya ('18)

Heinrich Göttlinger ('17)

Charles Grose ('18)

Marvin J. Grubman ('17)

Valery Z. Grdzelishvili ('19)

Haitao Guo ('18)

Ju-Tao Guo ('18)

Scott S. Halstead ('17)

Marie-Louise Hammarskjold ('19)

Michele Hardy ('18)

Richard Hardy ('18)

Edward W. Harhaj ('18)

Reuben Harris ('19)

Ronald Harty ('18)

Biao He ('19)

Bin He ('18)

Patrick Hearing ('18)

Mark Heise ('17)

Ekaterina E. Heldwein ('18)

Scott Hensley ('18)

Hans H. Hirsch ('19)

Vanessa Hirsch ('18)

John Hiscott ('19)

Brenda Hogue ('18)

Geoffrey H. Holm ('19)

Eddie Holmes ('18)

Fred Homa ('18)

Edward A. Hoover ('18)

Thomas Hope ('18)

Stacy M. Horner ('18)

Jianming Hu ('19)

Stephen Hughes ('18)

Ronald M. Iorio ('19)

Stuart Isaacs ('18)

Bertram Jacobs ('17)

Michael A. Jarvis ('19)

David C. Johnson ('17)

Marc C. Johnson ('19)

Welkin Johnson ('18)

Clinton Jones ('19)

Robert Kalejta ('18)

Jeremy Kamil ('19)

Stephanie M. Karst ('18)

Richard A. Katz ('19)

Yoshihiro Kawaoka ('19)

Kenneth Kaye ('18)

Brandon Keele ('18)

Shannon Kenney ('17)

Stephen J. Kent ('18)

Rajiv Khanna ('17)

Alexander Khromykh ('19)

Margaret Kielian ('18)

Paul R. Kinchington ('17)

William B. Klimstra ('17)

David Knipe ('17)

Richard A. Koup ('19)

Konstantin Kousoulas ('17)

Timothy F. Kowalik ('19)

Christine Kozak ('17)

Florian Krammer ('19)

Peter Krell ('18)

Thomas Krey ('19)

Thomas M. Kristie ('19)

Laurie Krug ('19)

Mark Krystal ('17)

Jens H. Kuhn ('17)

Richard Kuhn ('19)

Deanna Kulpa ('19)

Stefan Kunz ('17)

Michael Lai ('18)

Laimonis Laimins ('18)

Robert A. Lamb ('18)

Ke Lan ('17)

Nathaniel Landau ('18)

Ryan A. Langlois ('18)

Benhur Lee ('18)

David A. Leib ('17)

Keith Leppard ('18)

Laura Levy ('19)

Paul M. Lieberman ('17)

Brett Lindenbach ('19)

Maxine L. Linial ('19)

Howard Lipton ('18)

Shan-Lu Liu ('19)

Richard Lloyd ('18)

Dan Loeb ('19)

Volker Lohmann ('19)

Anice Lowen ('19)

Shan Lu ('19)

Jeremy Luban ('17)

Micah A. Luftig ('17)

Guangxiang G. Luo ('19)

Margaret MacDonald ('19)

Jason Mackenzie ('18)

Erich Mackow ('18)

Suresh Mahalingam ('18)

Renaud Mahieux ('19)

Shinji Makino ('19)

Frank Maldarelli ('18)

Balaji Manicassamy ('18)

Luis Martinez-Sobrido ('18)

James Maruniak ('18)

Paul S. Masters ('19)

Wendy Maury ('18)

Alison McBride ('18)

Sarah McDonald ('18)

Andrew Mehle ('17)

Xiang-Jin Meng ('19)

Janet E. Mertz ('19)

Thomas Mettenleiter ('17)

Craig Meyers ('19)

Cathy L. Miller ('18)

Chris Miller ('18)

Edward S. Mocarski ('17)

Ian Mohr ('19)

Ronald C. Montelaro ('17)

Nathaniel Moorman ('19)

Thomas Moran ('19)

Thomas E. Morrison ('17)

Trudy Morrison ('18)

Anne Moscona ('17)

Ashlee V. Moses ('17)

Bernard Moss ('19)

Karen L. Mossman ('17)

Walther Mothes ('18)

Karl Münger ('18)

Eain A. Murphy ('19)

Nicholas A. Muzyczka ('17)

Joe S. Mymryk ('19)

Peter Nagy ('19)

Stuart J. Neil ('19)

Glen R. Nemerow ('18)

Gabriele Neumann ('17)

Lisa F. P. Ng ('18)

Anthony V. Nicola ('18)

Jack Nunberg ('17)

Christine O'Connor ('19)

Peter O'Hare ('18)

Akira Ono ('19)

Klaus Osterrieder ('19)

Slobodan Paessler ('17)

Mirko Paiardini ('17)

Peter Palese ('19)

Massimo Palmarini ('18)

Leslie Parent ('19)

John Parker ('18)

Griffith D. Parks ('19)

Colin R. Parrish ('19)

A. Lorena Passarelli ('17)

Vinay K. Pathak ('17)

John Patton ('18)

William A. Paxton ('19)

Mark E. Peeples ('19)

Andrew Pekosz ('18)

Philip E. Pellett ('19)

Daniel Perez ('18)

Thomas Pietschmann ('17)

David J. Pintel ('19)

James M. Pipas ('17)

Vicente Planelles ('18)

Richard K. Plemper ('18)

Alexander Ploss ('19)

Eric Poeschla ('18)

Stefan H. Pöhlmann ('19)

B. V. Venkataram Prasad ('19)

Peter Prevelige ('18)

Jianming Qiu ('19)

Nancy Raab-Traub ('19)

Vincent R. Racaniello ('17)

Glenn Rall ('19)

Irene Ramos ('17)

Glenn Randall ('17)

Ranjit Ray ('18)

Stuart C. Ray ('19)

Matthias Reddehase ('18)

Barbara Rehermann ('19)

Alan R. Rein ('17)

Rolf Renne ('19)

Andrew Rice ('18)

Bert K. Rima ('19)

Michael Robek ('18)

Erle S. Robertson ('19)

Harriet L. Robinson ('17)

George Rohrmann ('19)

Bernard Roizman ('18)

Richard J. Roller ('17)

Monica Roth ('19)

Stefan Rothenburg ('17)

Raymond R. Rowland ('19)

Charles Russell ('19)

Anthony Sadler ('19)

Takeshi Saito ('19)

Jeffery T. Sample ('19)

Charles E. Samuel ('17)

R. Jude Samulski ('17)

Peter Sarnow ('17)

Quentin Sattentau ('18)

Daniel Sauter ('19)

Luis M. Schang ('19)

Matthias J. Schnell ('17)

John Schoggins ('18)

Herman B. Scholthof ('17)

Olivier Schwartz ('19)

Martin Schwemmle ('17)

Bert L. Semler ('19)

Megan L. Shaw ('19)

Pei-Yong Shi ('17)

Joanna L. Shisler ('17)

Sujan Shresta ('19)

Luis J. Sigal ('17)

Robert Siliciano ('18)

Viviana Simon ('19)

Alison Sinclair ('18)

John Sinclair ('19)

Mark Slifka ('19)

James Smiley ('18)

Greg Smith ('18)

Eric Snijder ('18)

Isabel Sola ('18)

Alicia Solorzano ('19)

Hugo Soudeyns ('18)

Paul Spearman ('18)

Samuel Speck ('18)

Deborah Spector ('18)

Katherine R. Spindler ('17)

Arun Srivastava ('18)

Leonidas Stamatatos ('18)

John Steel ('18)

Thilo Stehle ('18)

Jonathan P. Stoye ('17)

Klaus Strebel ('18)

William Sugden ('17)

Christopher S. Sullivan ('17)

Ren Sun ('19)

Mehul Suthar ('17)

Gerd Sutter ('17)

Nobuhiro Suzuki ('18)

Yuri Svitkin ('18)

Sankar Swaminathan ('19)

Ronald Swanstrom ('19)

Toru Takimoto ('17)

Vera L. Tarakanova ('19)

Michelle Tate ('18)

Peter Tattersall ('19)

John E. Tavis ('17)

John M. Taylor ('19)

Alice Telesnitsky ('17)

Markus Thali ('19)

Volker Thiel ('17)

Robert Thimme ('19)

Paul G. Thomas ('19)

Scott Tibbetts ('18)

Georgia D. Tomaras ('18)

Shuping Tong ('18)

Greg J. Towers ('19)

Paula Traktman ('19)

David C. Tscharke ('19)

Stephen Turner ('19)

Steven Varga ('18)

Subhash Vasudevan ('18)

Jeanmarie Verchot ('17)

Subhash C. Verma ('19)

Marco Vignuzzi ('18)

Volker M. Vogt ('17)

Veronika von Messling ('18)

Christopher M. Walker ('19)

Aiming Wang ('17)

David Wang ('19)

Fred C. Wang ('18)

Scott Weaver ('19)

Susan Weiss ('18)

Matthew D. Weitzman ('18)

Sandra Weller ('17)

Sean Whelan ('18)

Judith White ('18)

K. Andrew White ('18)

Anna Whitfield ('17)

Jason K. Whitmire ('19)

Michael Whitt ('18)

J. Lindsay Whitton ('19)

Reed B. Wickner ('18)

John V. Williams ('17)

John W. Wills ('19)

Duncan W. Wilson ('17)

Patrick Wilson ('18)

Christiane Wobus ('19)

Scott W. Wong ('19)

Yan Xiang ('17)

Otto Yang ('18)

Weiming Yuan ('19)

Yan Yuan ('19)

Andrew Yurochko ('19)

Xuming Zhang ('19)

Jincun Zhao ('19)

Chunfu Zheng ('17)

Zhi-Ming Zheng ('17)

Fanxiu Zhu ('19)

Michael Zwick ('18)

The Journal of Virology (ISSN 1098-5514), a publication of the American Society for Microbiology (ASM), 1752 N St., N.W., Washington, DC 20036-2904, is devoted to the dissemination of fundamental knowledge concerning viruses of bacteria, plants, and animals. Investigators are invited to submit reports of original research in all areas of basic virology, including biochemistry, biophysics, genetics, immunology, morphology, physiology, and pathogenesis and immunity.

Journal of Virology became an online-only journal with Volume 89, Issue 1 (January, 2015). The final print issue was Volume 88, Issue 24 (December 2014). Previous print ISSN: 0022-538X.

Instructions to Authors can be found at http://journalitas.asm.org/t/49545 and are updated throughout the year. The Journal of Virology is issued twice monthly, one volume per year. For subscription prices, contact the Subscriptions Unit, ASM (asmjournals{at}subscriptionoffice.com). Correspondence relating to reprint orders should be directed to ASM Author Services (ASM.AuthorServicesSupport{at}cenveo.com), and correspondence relating to disposition of submitted manuscripts, proofs, and general editorial matters should be directed to the Journals Department, American Society for Microbiology, 1752 N St., N.W., Washington, DC 20036-2904. Phone: (202) 737-3600 (see http://journals.asm.org/site/misc/staffcontacts.xhtml ). ASM policies for permissions, commercial reprints, and ePrints can be found at http://journals.asm.org/site/misc/reprints.xhtml .

CODEN: JOVIAM

doi:10.1128/JVI.masthead.91-10