Editor in Chief
Lynn W. Enquist (2012)
Princeton University

Editors
Karen L. Beemon (2012)
Johns Hopkins University

Byron W. Caughey (2013)
Rocky Mountain Laboratories

Terence S. Dermody (2009)
Vanderbilt University School of Medicine

Michael S. Diamond (2013)
Washington University School of Medicine

Daniel C. DiMaio, Cover Editor (2011)
Yale University School of Medicine

Robert W. Doms (2010)
University of Pennsylvania School of Medicine

Donald E. Ganem, Minireview Editor (2012)
University of California, San Francisco

Harry B. Greenberg (2013)
Stanford University School of Medicine

Beatrice H. Hahn (2009)
University of Alabama

Michael J. Imperiale (2011)
University of Michigan Medical School

Richard A. Koup (2013)
National Institute of Allergy and
Infectious Diseases

Douglas S. Lyles (2012)
Wake Forest University

Grant McFadden (2012)
University of Florida

Jay A. Nelson (2012)
Oregon Health and Science University

Peter M. Palese (2010)
Mount Sinai School of Medicine

Stanley Perlman (2012)
University of Iowa

Nancy Raab-Traub (2011)
University of North Carolina

Susan R. Ross (2012)
University of Pennsylvania

Rozanne M. Sandri-Goldin (2010)
University of California, Irvine

Bert L. Semler (2012)
University of California, Irvine

Ganes C. Sen (2013)
The Cleveland Clinic Foundation

Anne Simon (2012)
University of Maryland

Wesley I. Sundquist (2009)
University of Utah

Chairman, Publications Board
Thomas E. Shenk

Director, Journals
Barbara M. Goldman

Production Editor
Judith Nedrow

Assistant Production Editors
Julie Nachman
Karen Esplin

Editorial Board
Paul Ahlquist ('11)
Chris Aiken ('09)
Gillian Air ('09)
Lorraine Albritton ('10)
Antonio Alcami ('10)
Elliot J. Androphy ('11)
Carlos F. Arias ('09)
Eric Arts ('09)
Ann Arvin ('09)
Walter J. Atwood ('10)
Steve Bachenheimer ('09)
Joel Baines ('09)
Susan C. Baker ('11)
Dennis H. Bamford ('11)
Bruce W. Banfield ('10)
Lawrence Banks ('11)
Ralph Baric ('09)
Eric Barklis ('11)
Dan Barouch ('09)
Ralf Bartenschlager ('11)
David J. Barton ('11)
Chris Basler ('09)
Jeffrey M. Bergelson ('11)
Ben Berkhout ('09)
Ken Berns ('09)
Michael R. Betts ('10)
John A. Blaho ('10)
Keril Blight ('09)
Gary W. Blissard ('10)
David C. Bloom ('10)
Michele Bouloy ('09)
Michel Brahic ('09)
Wade Bresnahan ('11)
David Brian ('09)
Margo A. Brinton ('11)
William Britt ('09)
Chris Broder ('09)
Jay Brown ('11)
Ursula J. Buchholz ('11)
Michael Buchmeier ('09)
Dennis R. Burton ('10)
Frederic D. Bushman ('09)
Andrew Byrnes ('10)
Ellen D. Cahir-McFarland ('10)
Craig Cameron ('09)
Gabriella Campadelli-Fiume ('10)
Ann Campbell ('09)
Michael J. Carter ('11)
Roberto Cattaneo ('09)
Wen Chang ('11)
Bruce W. Chesebro ('09)
Paul Clapham ('10)
John M. Coffin ('09)
Jeffrey I. Cohen ('10)
Anamaris Colberg-Poley ('09)
Marc S. Collett ('10)
Peter Collins ('09)
Teresa Compton ('09)
Richard C. Condit ('10)
Karl-Klaus Conzelmann ('10)
Susan F. Cotmore ('10)
James Crowe ('09)
Bryan Cullen ('10)
James N. Culver ('10)
Blossom Damania ('10)
Nancy Davis ('09)
Andrew J. Davison ('10)
William O. Dawson ('10)
Juan Carlos de la Torre ('11)
Mark R. Denison ('11)
Cynthia A. Derdeyn ('10)
Prashant Desai ('09)
Ronald Desrosiers ('09)
Ulrich Desselberger ('09)
Dimiter Dimitrov ('09)
Shou Wei Ding ('11)
Dirk Dittmer ('10)
Valerian Dolja ('11)
Esteban Domingo ('09)
Philip R. Dormitzer ('10)
Theo Dreher ('11)
Jean D. Dubuisson ('11)
Jackie Dudley ('09)
Rebecca E. Dutch ('09)
Ellie Ehrenfeld ('10)
Maribeth Eiden ('09)
Roselyn J. Eisenberg ('09)
John H. Elder ('11)
Richard M. Elliott ('10)
Michael Emerman ('09)
Alan Engelman ('09)
Mariano Esteban ('10)
Roger Everett ('10)
Erik Falck-Pedersen ('10)
Hung Fan ('11)
Paul J. Farrell ('10)
Michael Farzan ('09)
Heinrich U. Feldmann ('11)
Ana Fernandez-Sesma ('09)
Jane Flint ('11)
Ron Fouchier ('10)
Genoveffa Franchini ('09)
Manuel Franco ('09)
Nigel Fraser ('09)
Eric O. Freed ('11)
Daved H. Fremont ('10)
Teryl Frey ('09)
Ilya Frolov ('10)
Klaus Frueh ('10)
Robert S. Fujinami ('11)
Dana H. Gabuzda ('11)
Michael Gale ('10)
Thomas Gallagher ('11)
Robert Garcea ('09)
J. Victor Garcia-Martinez ('10)
Adolfo Garcia-Sastre ('10)
Richard Gaynor ('09)
Adam P. Geballe ('10)
Chou-Zen Giam ('09)
Donald Gilden ('11)
Stephen P. Goff ('09)
Tatyana V. Golovkina ('11)
Francisco Gonzalez-Scarano ('08)
Felicia Goodrum ('10)
Heinrich Göttlinger ('11)
Patrick Green ('10)
Charles Grose ('09)
Linda Guarino ('09)
Nancy Haigwood ('09)
Otto A. Haller ('11)
Linda Hanley-Bowdoin ('10)
J. Marie Hardwick ('11)
Reuben Harris ('10)
Ronald Harty ('09)
Kim Hasenkrug ('09)
S. Diane Hayward ('10)
Patrick Hearing ('09)
Franz Heinz ('09)
Mark Heise ('11)
Ann B. Hill ('11)
Vanessa Hirsch ('09)
James M. Hogle ('09)
Brenda Hogue ('09)
Eddie Holmes ('09)
Fred Homa ('09)
Edward A. Hoover ('09)
Thomas Hope ('09)
Curt M. Horvath ('10)
Jim Hoxie ('09)
Jianming Hu ('10)
Stephen Hughes ('09)
Lindsey Hutt-Fletcher ('11)
Bertram Jacobs ('11)
Michael A. Jarvis ('10)
Ronald T. Javier ('10)
Kuan-Teh Jeang ('09)
David C. Johnson ('11)
R. Paul Johnson ('10)
Welkin Johnson ('09)
Paul Jolicoeur ('11)
Jae Jung ('09)
David Kabat ('11)
Cheng Kao ('09)
Michael Katze ('11)
Yoshihiro Kawaoka ('10)
Kenneth Kaye ('09)
George Kemble ('11)
Shannon Kenney ('11)
Vineet N. KewalRamani ('09)
Alexander Khromykh ('10)
Elliott Kieff ('11)
Margaret Kielian ('10)
Paul R. Kinchington ('11)
Frank Kirchhoff ('10)
Karla Kirkegaard ('11)
William B. Klimstra ('11)
David Knipe ('11)
Georg Kochs ('10)
Sarah Kopecky-Bromberg ('10)
Ulrich Koszinowski ('09)
Mark Krystal ('11)
Richard Kuhn ('10)
Laimonis Laimins ('09)
Michael Lairmore ('09)
Robert A. Lamb ('09)
Paul F. Lambert ('10)
Nathaniel Landau ('09)

Editorial Board
Benhur Lee ('09)
David A. Leib ('11)
Andrew Leigh Brown ('09)
Stanley M. Lemon ('10)
Laura Levy ('10)
Jeffrey Lifson ('10)
Brett Lindenbach ('10)
Maxine L. Linial ('10)
Howard Lipton ('09)
Richard Lloyd ('09)
Dan Loeb ('10)
Volker Lohmann ('10)
Richard Longnecker ('10)
Susana López Charretón ('09)
Anice Lowen ('10)
Jeremy Luban ('11)
Guangxiang G. Luo ('10)
Margaret MacDonald ('10)
Erich Mackow ('09)
Shinji Makino ('10)
David M. Margolis ('11)
Malcolm A. Martin ('11)
John Mascola ('10)
William S. Mason ('09)
Paul S. Masters ('10)
Thomas Mettenleiter ('11)
A. Dusty Miller ('09)
Chris Miller ('09)
Edward S. Mocarski ('11)
David Montefiori ('09)
Ronald C. Montelaro ('11)
Darius Moradpour ('09)
Elizabeth Moran ('11)
Thomas Moran ('10)
Trudy Morrison ('09)
Casey Morrow ('09)
Anne Moscona ('11)
Ashlee V. Moses ('11)
Don Mosier ('09)
Bernard Moss ('10)
Karen L. Mossman ('11)
Richard W. Moyer ('11)
Karl Münger ('09)
Nicholas A. Muzyczka ('11)
Peter Nagy ('10)
Glen R. Nemerow ('09)
Stuart T. Nichol ('10)
Janko Nikolich-Zugich ('10)
Edward G. Niles ('10)
Douglas Nixon ('09)
Isabel Suarez Novella ('10)
Don Nuss ('09)
Jennifer Nyborg ('09)
Peter O'Hare ('09)
David A. Ornelles ('11)
Klaus Osterrieder ('10)
James Ou ('09)
Ralph A. Pantophlet ('11)
Leslie Parent ('10)
Greg Pari ('09)
Griffith D. Parks ('10)
Colin R. Parrish ('10)
John Patton ('09)
Mark E. Peeples ('10)
Andrew Pekosz ('09)
Philip E. Pellett ('10)
Michael Perdue ('09)
Lenore Pereira ('09)
Daniel Perez ('09)
B. Matija Peterlin ('11)
Louis Picker ('09)
Theodore C. Pierson ('10)
Thomas Pietschmann ('11)
David J. Pintel ('10)
James M. Pipas ('11)
Paula Pitha-Rowe ('10)
B. V. Venkataram Prasad ('10)
Vinayaka R. Prasad ('10)
Chris Preston ('09)
Vincent R. Racaniello ('11)
Kathryn L. Radke ('11)
Glenn Rall ('10)
Robert F. Ramig ('10)
Lee Ratner ('09)
Stuart C. Ray ('10)
Matthias Reddehase ('09)
Barbara Rehermann ('10)
Alan R. Rein ('11)
Rolf Renne ('10)
Wolfgang Resch ('11)
Marilyn Resh ('10)
Felix A. Rey ('10)
Andrew Rice ('09)
Alan Rickinson ('11)
Bert K. Rima ('10)
Charles R. Rinaldo ('11)
Erle S. Robertson ('10)
Harriet L. Robinson ('11)
George Rohrmann ('10)
Bernard Roizman ('09)
Raymond P. Roos ('10)
John K. Rose ('10)
Monica Roth ('10)
Rob Ruigrok ('09)
Kathleen Rundell ('11)
Bill Ruyechan ('09)
Charles E. Samuel ('11)
R. Jude Samulski ('11)
Peter Sarnow ('11)
Quentin Sattentau ('09)
Dorothea L. Sawicki ('09)
Priscilla A. Schaffer ('11)
Brian Schaffhausen ('10)
John Schiller ('10)
Connie S. Schmaljohn ('10)
Matthias S. Schnell ('11)
Herman B. Scholthof ('11)
Paul Sharp ('09)
Megan L. Shaw ('10)
Pei-Yong Shi ('11)
Kunitada Shimotohno ('10)
Stuart Siddell ('09)
Aleem Siddiqui ('09)
Robert Siliciano ('09)
Robert Silverman ('09)
Guido Silvestri ('09)
Tony Simmons ('09)
John Sinclair ('10)
John Sixbey ('10)
Mark Slifka ('10)
James Smiley ('09)
Greg Smith ('09)
Eric Snijder ('09)
Joseph Sodroski ('09)
Richard R. Spaete ('10)
Paul Spearman ('09)
Samuel Speck ('09)
Deborah Spector ('09)
Katherine R. Spindler ('11)
Arun Srivastava ('09)
Leonidas Stamatatos ('09)
Mark F. Stinski ('10)
Victor Stollar ('09)
C. Martin Stoltzfus ('11)
Jonathan P. Stoye ('11)
Klaus Strebel ('09)
Daniel Streblow ('10)
William Sugden ('11)
Ren Sun ('11)
Gerd Sutter ('11)
Richard Sutton ('09)
Yuri Svitkin ('09)
Sankar Swaminathan ('10)
John M. Taylor ('10)
Alice Telesnitsky ('11)
Timothy Tellinghuisen ('10)
Heinz-Juergen Thiel ('09)
David A. Thorley-Lawson ('10)
Steven Tracy ('09)
Paula Traktman ('10)
Terrance Tumpey ('09)
Kenneth L. Tyler ('10)
Luis P. Villarreal ('09)
Volker M. Vogt ('11)
Bruce Walker ('09)
Christopher Walker ('09)
Fred C. Wang ('09)
Richard Ward ('09)
David Watkins ('11)
Scott Weaver ('10)
Susan Weiss ('09)
Sandra Weller ('11)
Judith White ('09)
J. Lindsay Whitton ('10)
Reed B. Wickner ('09)
Luc L. Willems ('11)
Bryan Williams ('10)
Carolyn Williamson ('10)
Scott W. Wong ('10)
David L. Woodland ('10)
Richard Wyatt ('11)
Otto Yang ('09)
T. S. Benedict Yen ('10)
Jonathan Yewdell ('11)
Jerome Zack ('10)