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Editorial Board

EDITOR IN CHIEF (2017)

Rozanne M. Sandri-Goldin

University of California, Irvine

EDITORS

Karen L. Beemon, Editor (2017) Johns Hopkins University

Byron W. Caughey, Editor (2013) Rocky Mountain Laboratories

Terence S. Dermody, Editor (2014) Vanderbilt University School of Medicine

Michael S. Diamond, Editor (2013) Washington University School of Medicine

Daniel C. DiMaio, Cover Editor (2016) Yale University School of Medicine

Robert W. Doms, Editor (2015) University of Pennsylvania School of Medicine

Klaus Frueh, Editor (2017) Oregon Health & Science University

Adolfo García-Sastre, Editor (2014) Mount Sinai School of Medicine

Stephen P. Goff, Minireview Editor (2015) Columbia University

Harry B. Greenberg, Editor (2013) Stanford University School of Medicine

Beatrice H. Hahn, Editor (2014) University of Pennsylvania

Lindsey Hutt-Fletcher, Editor (2014) Louisiana State University Health Science Center

Michael J. Imperiale, Editor (2016) University of Michigan Medical School

Karla Kirkegaard, Editor (2017) Stanford University

Richard A. Koup, Editor (2013) National Institute of Allergy and Infectious Diseases

Richard M. Longnecker, Editor (2016) Northwestern University

Douglas S. Lyles, Editor (2017) Wake Forest University

Grant McFadden, Editor (2017) University of Florida

Stanley Perlman, Editor (2017) University of Iowa

Susan R. Ross, Editor (2017) University of Pennsylvania

Ganes C. Sen, Editor (2013) The Cleveland Clinic Foundation

Anne Simon, Editor (2017) University of Maryland

Wesley I. Sundquist, Editor (2014) University of Utah

Thomas E. Shenk, Chairman, Publications Board

Barbara M. Goldman, Director, Journals

Judith Nedrow, Production Editor

Karen Esplin, Assistant Production Editor

Julie Nachman, Assistant Production Editor

EDITORIAL BOARD

Tero Ahola ('15)

Chris Aiken ('15)

Gillian Air ('15)

Randy Albrecht ('15)

Lorraine Albritton ('13)

Antonio Alcami ('13)

Louis A. Altamura ('13)

James Alwine ('15)

Elliot J. Androphy ('14)

Carlos F. Arias ('15)

Ann Arvin ('15)

Walter J. Atwood ('13)

Steve Bachenheimer ('15)

Joel Baines ('15)

Susan C. Baker ('14)

Dennis H. Bamford ('14)

Bruce W. Banfield ('13)

Lawrence Banks ('14)

Ralph Baric ('15)

Eric Barklis ('14)

Dan Barouch ('15)

Alan Barrett ('15)

Michele M. Barry ('13)

Ralf Bartenschlager ('14)

David J. Barton ('14)

Erik Barton ('15)

Jason C. Bartz ('14)

Chris Basler ('15)

George Belov ('15)

Jack R. Bennink ('13)

Jeffrey M. Bergelson ('14)

Cornelia C. Bergmann ('13)

Ben Berkhout ('15)

Kristen A. Bernard ('15)

Sonja Best ('15)

Michael R. Betts ('13)

James Binley ('15)

John A. Blaho ('13)

Joel Blankson ('15)

Gary W. Blissard ('13)

Jonas Blomberg ('15)

David C. Bloom ('13)

Kathleen Boris-Lawrie ('15)

Michele Bouloy ('15)

Michel Brahic ('15)

Jason Brenchley ('13)

Wade Bresnahan ('14)

David Brian ('15)

Margo A. Brinton ('14)

William Britt ('15)

Chris Broder ('15)

Jay Brown ('14)

Ursula J. Buchholz ('14)

Michael Buchmeier ('15)

Christopher B. Buck ('14)

Andrew Byrnes ('13)

Ellen D. Cahir-McFarland ('13)

Craig Cameron ('15)

Gabriella Campadelli-Fiume ('13)

Roberto Cattaneo ('15)

Wen Chang ('14)

Bruce W. Chesebro ('15)

Gregory Chinchar ('15)

Paul Clapham ('13)

Rollie Clem ('15)

Jeffrey I. Cohen ('13)

Anamaris Colberg-Poley ('15)

Peter Collins ('15)

Ronald Collman ('15)

Richard C. Condit ('13)

John H. Connor ('13)

Karl-Klaus Conzelmann ('13)

Susan F. Cotmore ('13)

Barbara S. Coulson ('13)

Carolyn Coyne ('15)

Rebecca Craven ('15)

Bryan Cullen ('13)

James N. Culver ('13)

Blossom Damania ('13)

Pranav Danthi ('13)

Andrew J. Davison ('13)

William O. Dawson ('13)

Victor DeFilippis ('15)

Juan Carlos de la Torre ('14)

Eric Delwart ('15)

Mark R. Denison ('14)

Cynthia A. Derdeyn ('13)

Prashant Desai ('15)

Ronald Desrosiers ('15)

Ulrich Desselberger ('15)

Paul Digard ('15)

Dimiter Dimitrov ('15)

Shou-Wei Ding ('14)

Dirk Dittmer ('13)

Valerian Dolja ('14)

Esteban Domingo ('15)

Philip R. Dormitzer ('13)

Christian Drosten ('15)

Jean D. Dubuisson ('14)

Jaquelin Dudley ('15)

Lynn B. Dustin ('13)

Rebecca E. Dutch ('15)

Roselyn J. Eisenberg ('15)

John H. Elder ('14)

Richard M. Elliott ('13)

Alan Engelman ('15)

Luis Enjuanes ('15)

Mariano Esteban ('13)

David Evans ('15)

Matthew J. Evans ('13)

Roger Everett ('13)

Erik Falck-Pedersen ('13)

Hung Fan ('14)

Bentley Fane ('15)

Paul J. Farrell ('13)

Michael Farzan ('15)

Heinrich U. Feldmann ('14)

Pinghui Feng ('15)

Ana Fernandez-Sesma ('15)

Jane Flint ('14)

Ron Fouchier ('13)

Manuel Franco ('15)

Lori Frappier ('13)

Brenda L. Fredericksen ('13)

Eric O. Freed ('14)

Ilya Frolov ('13)

Robert S. Fujinami ('14)

Dana H. Gabuzda ('14)

Michael Gale ('13)

Thomas Gallagher ('14)

Robert Garcea ('15)

Fernando Garcia-Arenal ('15)

Adam P. Geballe ('13)

Chou-Zen Giam ('15)

Donald Gilden ('14)

Paul Goepfert ('15)

Tatyana V. Golovkina ('14)

Felicia Goodrum ('13)

Alexander Gorbalenya ('15)

Heinrich Göttlinger ('14)

Barney Graham ('15)

Patrick Green ('13)

Charles Grose ('15)

Marvin J. Grubman ('14)

John Guatelli ('13)

Nancy Haigwood ('15)

Michele Hardy ('15)

Richard Hardy ('15)

Reuben Harris ('13)

Ronald Harty ('15)

S. Diane Hayward ('13)

Biao He ('13)

Bin He ('15)

Patrick Hearing ('15)

Mark Heise ('14)

Scott Hensley ('15)

Vanessa Hirsch ('15)

James M. Hogle ('15)

Brenda Hogue ('15)

Eddie Holmes ('15)

Fred Homa ('15)

Edward A. Hoover ('15)

Thomas Hope ('15)

Curt M. Horvath ('13)

Jianming Hu ('13)

Stephen Hughes ('15)

Ronald M. Iorio ('13)

Stuart Isaacs ('15)

Bertram Jacobs ('14)

Michael A. Jarvis ('13)

Kuan-Teh Jeang ('15)

David C. Johnson ('14)

R. Paul Johnson ('13)

Welkin Johnson ('15)

Kathryn Jones ('15)

Robert Kalejta ('15)

Stephanie M. Karst ('15)

Michael Katze ('14)

Yoshihiro Kawaoka ('13)

Michael S. Kay ('13)

Kenneth Kaye ('15)

Brandon Keele ('15)

George Kemble ('14)

Shannon Kenney ('14)

Stephen J. Kent ('15)

Alexander Khromykh ('13)

Elliott Kieff ('14)

Margaret Kielian ('15)

Paul R. Kinchington ('14)

Frank Kirchhoff ('13)

William B. Klimstra ('14)

David Knipe ('14)

Georg Kochs ('13)

Christine Kozak ('14)

Peter Krell ('15)

Mark Krystal ('14)

Richard Kuhn ('13)

Michael Lai ('15)

Laimonis Laimins ('15)

Robert A. Lamb ('15)

Paul F. Lambert ('13)

Nathaniel Landau ('15)

Benhur Lee ('15)

David A. Leib ('14)

Stanley M. Lemon ('13)

Keith Leppard ('15)

Laura Levy ('13)

Jeffrey Lifson ('13)

Brett Lindenbach ('13)

Maxine L. Linial ('13)

Howard Lipton ('15)

Richard Lloyd ('15)

Dan Loeb ('13)

Volker Lohmann ('13)

Susana López ('15)

Anice Lowen ('13)

Jeremy Luban ('14)

Guangxiang G. Luo ('13)

Margaret MacDonald ('13)

Erich Mackow ('15)

Shinji Makino ('13)

David M. Margolis ('14)

Luis Martinez-Sobrido ('15)

James Maruniak ('15)

John Mascola ('13)

William S. Mason ('15)

Paul S. Masters ('13)

Wendy Maury ('15)

Alison McBride ('15)

Sarah McDonald ('15)

Ilhem Messaoudi ('15)

Thomas Mettenleiter ('14)

A. Dusty Miller ('15)

Cathy L. Miller ('15)

Chris Miller ('15)

Edward S. Mocarski ('14)

Ian Mohr ('13)

David Montefiori ('15)

Ronald C. Montelaro ('14)

Thomas Moran ('13)

Lynn Morris ('13)

Trudy Morrison ('15)

Anne Moscona ('14)

Ashlee V. Moses ('14)

Don Mosier ('15)

Bernard Moss ('13)

Karen L. Mossman ('14)

Richard W. Moyer ('14)

Karl Münger ('15)

Nicholas A. Muzyczka ('14)

Peter Nagy ('13)

Stuart J. Neil ('13)

Glen R. Nemerow ('15)

Stuart T. Nichol ('13)

Anthony V. Nicola ('15)

Janko Nikolich-Zugich ('13)

Peter O'Hare ('15)

David A. Ornelles ('14)

Klaus Osterrieder ('13)

James Ou ('15)

Julie Overbaugh ('13)

Peter Palese ('13)

Massimo Palmarini ('15)

Ralph A. Pantophlet ('14)

Leslie Parent ('13)

Greg Pari ('15)

John Parker ('15)

Griffith D. Parks ('13)

Colin R. Parrish ('13)

Vinay K. Pathak ('14)

John Patton ('15)

William A. Paxton ('13)

Mark E. Peeples ('13)

Andrew Pekosz ('15)

Philip E. Pellett ('13)

Michael Perdue ('15)

Daniel Perez ('15)

Julie K. Pfeiffer ('15)

Theodore C. Pierson ('13)

Thomas Pietschmann ('14)

David J. Pintel ('13)

James M. Pipas ('14)

Vicente Planelles ('15)

Eric Poeschla ('15)

Stefan H. Pöhlmann ('13)

B. V. Venkataram Prasad ('13)

Peter Prevelige ('15)

Vincent R. Racaniello ('14)

Glenn Rall ('13)

Robert F. Ramig ('13)

Glenn Randall ('14)

Ranjit Ray ('15)

Stuart C. Ray ('13)

Matthias Reddehase ('15)

Barbara Rehermann ('13)

Alan R. Rein ('14)

Rolf Renne ('13)

Felix A. Rey ('13)

Andrew Rice ('15)

Alan Rickinson ('14)

Bert K. Rima ('13)

Michael Robek ('15)

Erle S. Robertson ('13)

Harriet L. Robinson ('14)

George Rohrmann ('13)

Bernard Roizman ('15)

Richard J. Roller ('14)

Monica Roth ('13)

Rob Ruigrok ('15)

Marco Salemi ('15)

Charles E. Samuel ('14)

R. Jude Samulski ('14)

Peter Sarnow ('14)

Quentin Sattentau ('15)

Dorothea L. Sawicki ('15)

Brian Schaffhausen ('13)

Connie S. Schmaljohn ('13)

Matthias J. Schnell ('14)

Herman B. Scholthof ('14)

Stacey Schultz-Cherry ('13)

Olivier Schwartz ('13)

Megan L. Shaw ('13)

Pei-Yong Shi ('14)

Aleem Siddiqui ('15)

Luis J. Sigal ('14)

Robert Siliciano ('15)

Guido Silvestri ('15)

Viviana Simon ('13)

Alison Sinclair ('15)

John Sinclair ('13)

Mark Slifka ('13)

James Smiley ('15)

Greg Smith ('15)

Eric Snijder ('15)

Paul Spearman ('15)

Samuel Speck ('15)

Deborah Spector ('15)

Katherine R. Spindler ('14)

Arun Srivastava ('15)

Leonidas Stamatatos ('15)

John Steel ('15)

Thilo Stehle ('15)

David A. Steinhauer ('13)

Jonathan P. Stoye ('14)

Klaus Strebel ('15)

Daniel Streblow ('13)

Kanta Subbarao ('13)

William Sugden ('14)

Christopher S. Sullivan ('14)

Nancy Sullivan ('14)

Gerd Sutter ('14)

Yuri Svitkin ('15)

Sankar Swaminathan ('13)

Ronald Swanstrom ('13)

John M. Taylor ('13)

Alice Telesnitsky ('14)

Timothy Tellinghuisen ('13)

Markus Thali ('13)

Volker Thiel ('14)

Scott Tibbetts ('15)

Georgia D. Tomaras ('15)

Shuping Tong ('15)

Greg J. Towers ('13)

Paula Traktman ('13)

Billy Tsai ('15)

David C. Tscharke ('13)

Steven Varga ('15)

Jeanmarie Verchot ('14)

Marco Vignuzzi ('15)

Volker M. Vogt ('14)

Veronika von Messling ('15)

Aiming Wang ('14)

David Wang ('13)

Fred C. Wang ('15)

David Watkins ('14)

Scott Weaver ('13)

Susan Weiss ('15)

Matt Weitzman ('15)

Sandra Weller ('14)

Raymond M. Welsh ('13)

Sean Whelan ('15)

Judith White ('15)

K. Andrew White ('15)

Jason K. Whitmire ('13)

Michael Whitt ('15)

J. Lindsay Whitton ('13)

Reed B. Wickner ('15)

Bryan Williams ('13)

John V. Williams ('14)

Duncan W. Wilson ('14)

Patrick Wilson ('15)

Scott W. Wong ('13)

Otto Yang ('15)

Dong Yu ('13)

Xiaofang Yu ('13)

Andrew Yurochko ('13)

Jerome Zack ('13)

Michael Zwick ('15)

The Journal of Virology (ISSN 0022-538X), a publication of the American Society for Microbiology (ASM), 1752 N St., N.W., Washington, DC 20036-2904, is devoted to the dissemination of fundamental knowledge concerning viruses of bacteria, plants, and animals. Investigators are invited to submit reports of original research in all areas of basic virology, including biochemistry, biophysics, genetics, immunology, morphology, physiology, and pathogenesis and immunity. Instructions to authors are no longer published in the first issue of the year but can be found at http://jvi.asm.org/site/misc/ifora.xhtml and are updated throughout the year. The Journal is issued twice monthly, one volume per year. Nonmember print subscription prices (per year) are: $2,545, U.S.; $2,736, Canada (plus 13% HST where applicable); $2,743, Europe; $2,810, Latin America; $2,818, rest of world. Member print subscription prices (per year) are: $537, U.S.; $557, Canada (plus 13% HST where applicable); $564, Europe; $646, Latin America; $637, rest of world. Single copies are: $100, nonmember; $50, member (Canadians add 6% GST, or 14% GST+HST where applicable). For prices of electronic versions, contact the Subscriptions Unit, ASM. Correspondence relating to subscriptions, defective copies, missing issues, and availability of back issues should be directed to the Subscriptions Unit, ASM; correspondence relating to reprint orders should be directed to the Reprint Order Unit, ASM; and correspondence relating to disposition of submitted manuscripts, proofs, and general editorial matters should be directed to the Journals Department, American Society for Microbiology, 1752 N St., N.W., Washington, DC 20036-2904. Phone: (202) 737-3600.

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doi:10.1128/JVI.masthead.87-3

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